故事背景为1937年秋天，隶属於日本帝国的的结城中校不顾陆军高层反弹，於军队中成立了间谍培训学校“D机关”。结城教导这些刚毕业自各地大学的年轻人各种军事能力，并灌输所有成员否定军人三大戒律“不准死”、“不准杀人”及“不准被抓”的思想。来自东京、上海和伦敦等地的各国间谍展开一连串发挥各自谋略和头脑，甚至不惜欺骗友军的生死游戏。
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